About the Authors

David C. Kazyak

dkazyak@gmail.com

Affiliations University of Maryland Center for Environmental Science, Appalachian Laboratory, Frostburg, Maryland, United States of America, U.S. Geological Survey, Leetown Science Center, Aquatic Ecology Branch, Kearneysville, West Virginia, United States of America

Robert H. Hilderbrand

Affiliation University of Maryland Center for Environmental Science, Appalachian Laboratory, Frostburg, Maryland, United States of America

Tim L. King

Affiliation U.S. Geological Survey, Leetown Science Center, Aquatic Ecology Branch, Kearneysville, West Virginia, United States of America

Stephen R. Keller

Affiliation University of Vermont, Department of Plant Biology, Burlington, Vermont, United States of America

Vikram E. Chhatre

Affiliation University of Vermont, Department of Plant Biology, Burlington, Vermont, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DCK RHH TLK SRK VEC. Performed the experiments: DCK RHH TLK. Analyzed the data: DCK RHH TLK SRK VEC. Contributed reagents/materials/analysis tools: DCK RHH TLK. Wrote the paper: DCK RHH TLK SRK VEC.