About the Authors

Kyle A. Garver

Contributed equally to this work with: Kyle A. Garver, Stewart C. Johnson

Kyle.Garver@dfo-mpo.gc.ca

Affiliation Pacific Biological Station, Department of Fisheries and Oceans, Nanaimo, British Columbia, Canada

Stewart C. Johnson

Contributed equally to this work with: Kyle A. Garver, Stewart C. Johnson

Affiliation Pacific Biological Station, Department of Fisheries and Oceans, Nanaimo, British Columbia, Canada

Mark P. Polinski

Affiliation Pacific Biological Station, Department of Fisheries and Oceans, Nanaimo, British Columbia, Canada

Julia C. Bradshaw

Affiliation Pacific Biological Station, Department of Fisheries and Oceans, Nanaimo, British Columbia, Canada

Gary D. Marty

Affiliation Animal Health Centre, Ministry of Agriculture, Abbotsford, British Columbia, Canada

Heindrich N. Snyman

Affiliation Animal Health Centre, Ministry of Agriculture, Abbotsford, British Columbia, Canada

Diane B. Morrison

Affiliation Marine Harvest Canada, Campbell River, British Columbia, Canada

Jon Richard

Affiliation Pacific Biological Station, Department of Fisheries and Oceans, Nanaimo, British Columbia, Canada

Competing Interests

Partial funding for this study was provided by Marine Harvest Canada. This commercial affiliation does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KAG SCJ. Performed the experiments: KAG SCJ MPP JCB JR. Analyzed the data: KAG SCJ MPP JCB GDM HNS. Contributed reagents/materials/analysis tools: DBM. Wrote the paper: KAG MPP SCJ GDM. Conducted histological examinations: GDM HNS.