About the Authors

Kwang-Hwan Kim

Affiliation Department of Horticultural Bioscience, Pusan National University, Miryang, Republic of Korea

Ji-Hyun Hwang

Affiliation Department of Horticultural Bioscience, Pusan National University, Miryang, Republic of Korea

Dong-Yeup Han

Affiliation Department of Horticultural Bioscience, Pusan National University, Miryang, Republic of Korea

Minkyu Park

Affiliations Department of Plant Science, Seoul National University, Seoul, Republic of Korea, Department of Genetics, University of Georgia, Athens, Georgia, United States of America

Seungill Kim

Affiliation Department of Plant Science, Seoul National University, Seoul, Republic of Korea

Doil Choi

Affiliation Department of Plant Science, Seoul National University, Seoul, Republic of Korea

Yongjae Kim

Affiliation Partner Seed, Ansung, Republic of Korea

Gung Pyo Lee

Affiliation Department of Integrative Plant Science, College of Biotechnology and Natural Resources, Chung-Ang University, Ansung, Republic of Korea

Sun-Tae Kim

Affiliation Department of Plant Bioscience, Pusan National University, Miryang, Republic of Korea

Young-Hoon Park

ypark@pusan.ac.kr

Affiliations Department of Horticultural Bioscience, Pusan National University, Miryang, Republic of Korea, Life and Industry Convergence Research Institute, Pusan National University, Miryang, Republic Korea

Competing Interests

The authors have declared that no competing interests exist. A commercial affiliation, "Partner Seed" (affiliated author: Yongjae Kim) did not provide financial support, but played a role in the study design and data collection by field evaluation of the F2 population. Phenotyping of the F2 population was conducted by Yongjae Kim (the owner of "Partner seed") at the breeding field owned by Partner Seed Company which collaborated for this study. Partner Seed agreed to disclose any information described in this manuscript under no condition. Therefore, this does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KHK DC YK GPL STK YP. Performed the experiments: KHK JHH DYH MP SK YK YP. Analyzed the data: KHK DYH MP SK YP. Contributed reagents/materials/analysis tools: KHK MP SK DC YK YP. Wrote the paper: KHK DYH MP SK YP.