About the Authors

Ushio Minami

Contributed equally to this work with: Ushio Minami, Mariko Nishi, Taro Fukaya, Yoshinori Fujiwara

minami@tmig.or.jp

Affiliations Research Team for Social Participation and Community Health, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan, Graduate School of Human Science and Environment, University of Hyogo, Hyogo, Japan

Mariko Nishi

Contributed equally to this work with: Ushio Minami, Mariko Nishi, Taro Fukaya, Yoshinori Fujiwara

Affiliation Research Team for Social Participation and Community Health, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan

Taro Fukaya

Contributed equally to this work with: Ushio Minami, Mariko Nishi, Taro Fukaya, Yoshinori Fujiwara

Affiliation Department of Translational Research Promotion, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan

Masami Hasebe

‡ These authors also contributed equally to this work.

Affiliations Research Team for Social Participation and Community Health, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan, Faculty of Human Welfare, Seigakuin University, Saitama, Japan

Kumiko Nonaka

‡ These authors also contributed equally to this work.

Affiliation Research Team for Social Participation and Community Health, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan

Takashi Koike

‡ These authors also contributed equally to this work.

Affiliations Research Team for Social Participation and Community Health, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan, College of Humanities and Sciences, Nihon University, Tokyo, Japan

Hiroyuki Suzuki

‡ These authors also contributed equally to this work.

Affiliation Research Team for Social Participation and Community Health, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan

Yoh Murayama

‡ These authors also contributed equally to this work.

Affiliation Research Team for Social Participation and Community Health, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan

Hayato Uchida

‡ These authors also contributed equally to this work.

Affiliation Graduate School of Human Science and Environment, University of Hyogo, Hyogo, Japan

Yoshinori Fujiwara

Contributed equally to this work with: Ushio Minami, Mariko Nishi, Taro Fukaya, Yoshinori Fujiwara

Affiliation Research Team for Social Participation and Community Health, Tokyo Metropolitan Institute of Gerontology, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: UM YF MN TF HS. Performed the experiments: MN TF MH KN TK YM. Analyzed the data: UM HU YF. Contributed reagents/materials/analysis tools: UM TF MH HS YF. Wrote the paper: UM.