About the Authors

David M. Theobald

davet@csp-inc.org

Affiliations Conservation Science Partners, Truckee, California, United States of America, Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America, Graduate Degree Program in Ecology, Colorado State University, Fort Collins, Colorado, United States of America

Dylan Harrison-Atlas

Affiliation Graduate Degree Program in Ecology, Colorado State University, Fort Collins, Colorado, United States of America

William B. Monahan

Affiliation Inventory and Monitoring Division, National Park Service, Fort Collins, Colorado, United States of America

Christine M. Albano

Affiliations Conservation Science Partners, Truckee, California, United States of America, John Muir Institute of the Environment, University of California Davis, Davis, California, United States of America

Competing Interests

The authors of this manuscript have the following competing interests: David M. Theobald, Dylan Harrison-Atlas, and Christine M. Albano are employed by Conservation Science Partners. There are no patents, products in development, or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: DMT DH-A WBM CMA. Analyzed the data: DMT DH-A WBM. Wrote the paper: DMT DH-A WBM CMA.