About the Authors

Mary H. Straub

Affiliation Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Terra R. Kelly

Affiliation Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Bruce A. Rideout

Affiliation Wildlife Disease Laboratories, San Diego Zoo Institute for Conservation Research, Escondido, California, United States of America

Curtis Eng

Current address: Western University College of Veterinary Medicine, Pomona, California, United Stated of America

Affiliation Los Angeles Zoo and Botanical Gardens, Los Angeles, California, United States of America

Janna Wynne

Affiliation California Science Center Foundation, Los Angeles, California, United States of America

Josephine Braun

Affiliation Wildlife Disease Laboratories, San Diego Zoo Institute for Conservation Research, Escondido, California, United States of America

Christine K. Johnson

ckjohnson@ucdavis.edu

Affiliation Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MHS TRK CKJ BAR JB CE JW. Performed the experiments: BAR JB MHS. Analyzed the data: MHS TRK CKJ. Contributed reagents/materials/analysis tools: CE BAR JW. Wrote the paper: MHS TRK CKJ JB JW BAR CE.