About the Authors

Samantha J. Hau

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Jisun Sun

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, St Paul, Minnesota, United States of America

Peter R. Davies

Affiliation Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, St Paul, Minnesota, United States of America

Timothy S. Frana

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Tracy L. Nicholson

tracy.nicholson@ars.usda.gov

Affiliation National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Competing Interests

The Competing Interests include funding from a commercial source: The National Pork Board. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SJH JS PRD TSF TLN. Performed the experiments: SJH. Analyzed the data: SJH JS PRD TSF TLN. Contributed reagents/materials/analysis tools: SJH JS PRD TSF TLN. Wrote the paper: SJH JS PRD TSF TLN.