About the Authors

Eric Buffetaut

eric.buffetaut@sfr.fr

Affiliation Laboratoire de Géologie de l’Ecole Normale Supérieure, CNRS (UMR 8538), Paris, France

Axel-Frans Hartman

Affiliations Petroleum Development Oman, Muscat, Sultanate of Oman, Shell International Exploration and Production B.V, The Hague, the Netherlands

Mohammed Al-Kindi

Affiliation Petroleum Development Oman, Muscat, Sultanate of Oman

Anne S. Schulp

Affiliations Naturalis Biodiversity Center, Leiden, the Netherlands, Natuurhistorisch Museum Maastricht, Maastricht, the Netherlands, Faculty of Earth and Life Sciences, Amsterdam VU University, Amsterdam, the Netherlands

Competing Interests

The authors have the following interests: Mohammed al-Kindi is employed by Petroleum Development Oman. Axel F. Hartman is employed by Petroleum Development Oman and Shell International Exploration and Production B.V. Neither Petroleum Development Oman or Shell International Exploration and Production B.V. have been involved in this study. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Analyzed the data: EB AFH MK ASS. Wrote the paper: EB AFH MK ASS. Specimen collection from 2011-2014 in the area of Al Khod, Muscat in Oman: AFH.