About the Authors

Masanori Kohda

kohda.tanganyika@gmail.com

Affiliation Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Sciences, Osaka City University, Sumiyoshi, Osaka 558–8585, Japan

Lyndon Alexander Jordan

Current address: Integrative Biology, The University of Texas at Austin, Austin, Texas, United States of America

Affiliation Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Sciences, Osaka City University, Sumiyoshi, Osaka 558–8585, Japan

Takashi Hotta

Affiliation Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Sciences, Osaka City University, Sumiyoshi, Osaka 558–8585, Japan

Naoya Kosaka

Affiliation Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Sciences, Osaka City University, Sumiyoshi, Osaka 558–8585, Japan

Kenji Karino

Affiliation Department of Biology, Tokyo Gakugei University, Koganei, Tokyo 184–8501, Japan

Hirokazu Tanaka

Affiliation Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Sciences, Osaka City University, Sumiyoshi, Osaka 558–8585, Japan

Masami Taniyama

Affiliation Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Sciences, Osaka City University, Sumiyoshi, Osaka 558–8585, Japan

Tomohiro Takeyama

Current address: Department of Biosphere-Geosphere Science, Faculty of Biosphere-Geosphere Science, Okayama University of Science, Okayama 700–0005, Japan

Affiliation Laboratory of Animal Sociology, Department of Biology and Geosciences, Graduate School of Sciences, Osaka City University, Sumiyoshi, Osaka 558–8585, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MK KK TT. Performed the experiments: NK MT HT TH. Analyzed the data: LAJ TT HT. Contributed reagents/materials/analysis tools: HT TH. Wrote the paper: MK LAJ TT.