About the Authors

Bernardo Vargas-Ángel

Bernado.VargasAngel@noaa.gov

Affiliation Joint Institute for Marine and Atmospheric Research, University of Hawaii, Honolulu, Hawaii, 96818, United States of America

Cristi L. Richards

Affiliation 2525 Date St. Apt. 3101, Honolulu, Hawaii, 96826–5420, United States of America

Peter S. Vroom

Affiliation Ocean Associates, 1846 Wasp Blvd. Bldg., # 176, Honolulu, Hawaii, 96818, United States of America

Nichole N. Price

Affiliation Bigelow Laboratory for Ocean Sciences, 60 Bigelow Dr., East Boothbay, Maine, 04544, United States of America

Tom Schils

Affiliation University of Guam Marine Laboratory, Mangilao, Guam, 96913, United States of America

Charles W. Young

Affiliation Joint Institute for Marine and Atmospheric Research, University of Hawaii, Honolulu, Hawaii, 96818, United States of America

Jennifer Smith

Affiliation Scripps Institution of Oceanography, University of California San Diego, 9500 Gilman Dr., La Jolla, California, 92093, United States of America

Maggie D. Johnson

Affiliation Scripps Institution of Oceanography, University of California San Diego, 9500 Gilman Dr., La Jolla, California, 92093, United States of America

Russell E. Brainard

Affiliation NOAA Pacific Islands Fisheries Science Center, Coral Reef Ecosystem Division, 1846 Wasp Blvd. Bldg. # 176, Honolulu, Hawaii, 96818, United States of America

Competing Interests

Peter S. Vroom was employed by Ocean Associates at the time of manuscript preparation. There are no patents, products in development or marketed products to declare. This did not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: REB NNP JS CWY. Performed the experiments: CWY CLR. Analyzed the data: BVA PSV CLR NNP TS MDJ. Contributed reagents/materials/analysis tools: JS. Wrote the paper: BVA CLR PSV NNP TS MDJ REB.