About the Authors

Alistair M. McInnes

amcinnes3@gmail.com

Affiliation Percy FitzPatrick Institute, DST/NRF Centre of Excellence, University of Cape Town, Cape Town, South Africa

Arjun Khoosal

Affiliation Computational Biology Group, Institute of Infectious Diseases and Molecular Medicine, University of Cape Town, Cape Town, South Africa

Ben Murrell

Affiliation Department of Medicine, University of California San Diego, La Jolla, California, United States of America

Dagmar Merkle

Affiliation Branch: Fisheries Management, Department of Agriculture, Forestry and Fisheries, Rogge Bay, South Africa

Miguel Lacerda

Affiliation Department of Statistical Sciences, University of Cape Town, Cape Town, South Africa

Reason Nyengera

Affiliation DST/NRF Centre of Excellence at the Percy FitzPatrick Institute of African Ornithology, Department of Zoology, Nelson Mandela Metropolitan University, Port Elizabeth, South Africa

Janet C. Coetzee

Affiliation Branch: Fisheries Management, Department of Agriculture, Forestry and Fisheries, Rogge Bay, South Africa

Loyd C. Edwards

Affiliation Raggy Charters, Port Elizabeth, South Africa

Peter G. Ryan

Affiliation Percy FitzPatrick Institute, DST/NRF Centre of Excellence, University of Cape Town, Cape Town, South Africa

Johan Rademan

Affiliation Branch: Fisheries Management, Department of Agriculture, Forestry and Fisheries, Rogge Bay, South Africa

Jan J van der Westhuizen

Affiliation Branch: Fisheries Management, Department of Agriculture, Forestry and Fisheries, Rogge Bay, South Africa

Lorien Pichegru

Affiliation DST/NRF Centre of Excellence at the Percy FitzPatrick Institute of African Ornithology, Department of Zoology, Nelson Mandela Metropolitan University, Port Elizabeth, South Africa

Competing Interests

The authors have the following interests: Co-author Loyd C. Edwards is employed by Raggy Charters. There are no patents, products in development or marketed products to declare. This does not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AMM AK BM JCC PGR JR JJV LP. Performed the experiments: AMM DM RN LCE JR JJV LP. Analyzed the data: AMM AK BM ML JCC. Contributed reagents/materials/analysis tools: AK BM DM JCC. Wrote the paper: AMM BM ML JCC PGR LP. Designed software FISH for hydoacoustic quantification: AK BM.