About the Authors

Wayne T. A. Enanoria

Affiliation Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America

Lee Worden

Affiliation Francis I. Proctor Foundation for Research in Ophthalmology, University of California San Francisco, San Francisco, California, United States of America

Fengchen Liu

Affiliation Francis I. Proctor Foundation for Research in Ophthalmology, University of California San Francisco, San Francisco, California, United States of America

Daozhou Gao

Affiliation Francis I. Proctor Foundation for Research in Ophthalmology, University of California San Francisco, San Francisco, California, United States of America

Sarah Ackley

Affiliation Francis I. Proctor Foundation for Research in Ophthalmology, University of California San Francisco, San Francisco, California, United States of America

James Scott

Affiliations Francis I. Proctor Foundation for Research in Ophthalmology, University of California San Francisco, San Francisco, California, United States of America, Mathematics and Statistics, Colby College, Waterville, Maine, United States of America

Michael Deiner

Affiliation Department of Ophthalmology, University of California San Francisco, San Francisco, California, United States of America

Ernest Mwebaze

Affiliation Makerere University, Kampala, Uganda

Wui Ip

Affiliation Francis I. Proctor Foundation for Research in Ophthalmology, University of California San Francisco, San Francisco, California, United States of America

Thomas M. Lietman

Affiliations Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America, Francis I. Proctor Foundation for Research in Ophthalmology, University of California San Francisco, San Francisco, California, United States of America, Department of Ophthalmology, University of California San Francisco, San Francisco, California, United States of America

Travis C. Porco

travis.porco@ucsf.edu

Affiliations Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America, Francis I. Proctor Foundation for Research in Ophthalmology, University of California San Francisco, San Francisco, California, United States of America, Department of Ophthalmology, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WTAE TML TCP. Performed the experiments: TCP. Analyzed the data: WTAE TCP. Contributed reagents/materials/analysis tools: FL SA LW. Wrote the paper: WTAE LW FL DZG SA JS MD EM WI TML TCP.