About the Authors

Juliana Houghton

stephj5@uw.edu

Affiliation School of Aquatic & Fishery Sciences, University of Washington, Seattle, WA, United States of America

Marla M. Holt

Affiliation Conservation Biology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, Seattle, WA, United States of America

Deborah A. Giles

Affiliation Wildlife, Fish, & Conservation Biology, University of California Davis, Davis, CA, United States of America

M. Bradley Hanson

Affiliation Conservation Biology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, Seattle, WA, United States of America

Candice K. Emmons

Affiliation Conservation Biology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, Seattle, WA, United States of America

Jeffrey T. Hogan

Affiliation Cascadia Research Collective, Olympia, WA, United States of America

Trevor A. Branch

Affiliation School of Aquatic & Fishery Sciences, University of Washington, Seattle, WA, United States of America

Glenn R. VanBlaricom

Affiliations U.S. Geological Survey, Washington Cooperative Fish and Wildlife Research Unit, Seattle, WA, United States of America, School of Aquatic & Fishery Sciences, University of Washington, Seattle, WA, United States of America

Competing Interests

The authors of this manuscript have the following competing interests: MBH is partially responsible for funding decisions from Northwest Fisheries Science Center. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JH MMH MBH. Performed the experiments: JH MMH DAG MBH CKE JTH. Analyzed the data: JH MMH DAG TAB. Contributed reagents/materials/analysis tools: JH MMH DAG MBH. Wrote the paper: JH MMH MBH TAB GRV.