About the Authors

Anil Raj

Contributed equally to this work with: Anil Raj, Heejung Shim

rajanil@stanford.edu (AR); hjshim@gmail.com (HS); gilad@uchicago.edu (YG); pritch@stanford.edu (JKP); mstephens@uchicago.edu (MS)

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Heejung Shim

Contributed equally to this work with: Anil Raj, Heejung Shim

rajanil@stanford.edu (AR); hjshim@gmail.com (HS); gilad@uchicago.edu (YG); pritch@stanford.edu (JKP); mstephens@uchicago.edu (MS)

Current address: Department of Statistics, Purdue University, West Lafayette, Indiana, United States of America

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Yoav Gilad

rajanil@stanford.edu (AR); hjshim@gmail.com (HS); gilad@uchicago.edu (YG); pritch@stanford.edu (JKP); mstephens@uchicago.edu (MS)

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Jonathan K. Pritchard

rajanil@stanford.edu (AR); hjshim@gmail.com (HS); gilad@uchicago.edu (YG); pritch@stanford.edu (JKP); mstephens@uchicago.edu (MS)

Affiliations Department of Genetics, Stanford University, Stanford, California, United States of America, Department of Biology, Stanford University, Stanford, California, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Matthew Stephens

rajanil@stanford.edu (AR); hjshim@gmail.com (HS); gilad@uchicago.edu (YG); pritch@stanford.edu (JKP); mstephens@uchicago.edu (MS)

Affiliations Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America, Department of Statistics, University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AR HS YG JKP MS. Analyzed the data: AR HS. Wrote the paper: AR HS YG JKP MS.