About the Authors

Rowena M. A. Packer

rpacker@rvc.ac.uk

Affiliation Department of Clinical Science and Services, Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Anke Hendricks

Affiliation Department of Clinical Science and Services, Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Michael S. Tivers

Affiliation School of Veterinary Sciences, University of Bristol, Langford House, Langford, Bristol, BS40 5DU, United Kingdom

Charlotte C. Burn

Affiliation Department of Production and Population Health, Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Competing Interests

Michael S. Tivers was employed by Cave Veterinary Specialists during part of this study. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: RMAP AH CCB. Performed the experiments: RMAP MST. Analyzed the data: RMAP CCB. Contributed reagents/materials/analysis tools: RMAP MST. Wrote the paper: RMAP AH MST CCB. Diagnosed clinical cases: MST.