About the Authors

Hubiao Yang

Contributed equally to this work with: Hubiao Yang, Xiaoxia Li

Affiliation Tropical Crops Genetic Resources Institute, Chinese Academy of Tropic Agricultural Sciences, Danzhou, Hainan, People's Republic of China

Xiaoxia Li

Contributed equally to this work with: Hubiao Yang, Xiaoxia Li

Affiliation Environment and Plant Protection Institute, Chinese Academy of Tropical Agricultural Sciences, Haikou, Hainan, People's Republic of China

Wenqiang Wang

Affiliation Tropical Crops Genetic Resources Institute, Chinese Academy of Tropic Agricultural Sciences, Danzhou, Hainan, People's Republic of China

Changjun Bai

Affiliation Tropical Crops Genetic Resources Institute, Chinese Academy of Tropic Agricultural Sciences, Danzhou, Hainan, People's Republic of China

Guodao Liu

liuguodao2008@163.com

Affiliation Tropical Crops Genetic Resources Institute, Chinese Academy of Tropic Agricultural Sciences, Danzhou, Hainan, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HY XL. Performed the experiments: XL HY. Analyzed the data: HY XL. Contributed reagents/materials/analysis tools: GL CB WW. Wrote the paper: HY XL.