About the Authors

Linyuan Shen

Contributed equally to this work with: Linyuan Shen, Jia Luo

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Jia Luo

Contributed equally to this work with: Linyuan Shen, Jia Luo

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Jingjing Du

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Chendong Liu

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Xiaoqian Wu

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Qiang Pu

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Yuhua Fu

Affiliation Key Laboratory of Agricultural Animal Genetics, Breeding, and Reproduction of Ministry of Education & Key Laboratory of Swine Genetics, Huazhong Agricultural University, Wuhan, China

Qianzi Tang

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Yuanrui Liu

Affiliation LC Sciences, Hangzhou, Zhejiang, China

Qiang Li

Affiliation Sichuan Province General Station of Animal Husbandry, Chengdu, Sichuan, China

Runlin Yang

Affiliations College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China, Mabian Gold Liangshan Agricultural Development Co., Ltd, Sichuan, China

Xuewei Li

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Guoqing Tang

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Yanzhi Jiang

Affiliation Department of Biology, College of Life and Science, Sichuan Agricultural University, Chengdu, Sichuan, China

Mingzhou Li

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Shunhua Zhang

zhangsh1919@163.com (SZ); zhuli7508@163.com (LZ)

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Li Zhu

zhangsh1919@163.com (SZ); zhuli7508@163.com (LZ)

Affiliation College of Animal Science and Technology, Sichuan Agricultural University, Chengdu, Sichuan, China

Competing Interests

Runlin Yang was employed by Mabian Gold Liangshan Agricultural Development Co., Ltd. for part of this study and Yuanrui Liu by LC Sciences. There are no patents, products in development, or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: LYS LZ XWL SHZ. Performed the experiments: LYS RLY JJD CDL XQW QP QL GQT YZJ. Analyzed the data: LYS JL QZT YRL MZL YHF. Contributed reagents/materials/analysis tools: LYS RLY JL. Wrote the paper: LYS JL LZ. All authors read and approved the final manuscript.