About the Authors

Brad Seely

brad.seely@ubc.ca

Affiliation Department of Forest Resources Management, University of British Columbia, Vancouver, British Columbia, Canada

Clive Welham

Affiliation Department of Forest Resources Management, University of British Columbia, Vancouver, British Columbia, Canada

Kim Scoullar

Affiliation Life Sciences Programming Incorporated, Naramata, British Columbia, Canada

Competing Interests

One of the authors (KS) is employed by Life Sciences Programming, Inc. This does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: BS CW KS. Performed the experiments: BS. Analyzed the data: BS CW. Contributed reagents/materials/analysis tools: KS. Wrote the paper: BS CW KS. Code development: KS BS.