About the Authors

Susann K. J. Ludwig

s.k.j.ludwig@tue.nl

Current address: Laboratory of Chemical Biology, Department of Biomedical Engineering, Eindhoven University of Technology, Eindhoven, The Netherlands

Affiliation RIKILT Wageningen UR, Wageningen, The Netherlands

Christian Tokarski

Affiliation Biologisch-Pharmazeutische Fakultät, Friedrich-Schiller-Universität Jena, Jena, Germany

Stefan N. Lang

Affiliation Biologisch-Pharmazeutische Fakultät, Friedrich-Schiller-Universität Jena, Jena, Germany

Leendert A. van Ginkel

Affiliation RIKILT Wageningen UR, Wageningen, The Netherlands

Hongying Zhu

Affiliation Electrical Engineering Department, University of California Los Angeles, Los Angeles, California, United States of America

Aydogan Ozcan

Affiliations Electrical Engineering Department, University of California Los Angeles, Los Angeles, California, United States of America, Bioengineering Department, University of California Los Angeles, Los Angeles, California, United States America, California NanoSystems Institute (CNSI), University of California Los Angeles, Los Angeles, California, United States of America

Michel W. F. Nielen

Affiliations RIKILT Wageningen UR, Wageningen, The Netherlands, Laboratory of Organic Chemistry, Wageningen University, Wageningen, The Netherlands

Competing Interests

A. Ozcan is the co-founder of a start-up company (Holomic LLC) that commercializes computational microscopy and diagnostic tools. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: SKJL LG HZ AO MN. Performed the experiments: SKJL. Analyzed the data: SKJL CT SL. Contributed reagents/materials/analysis tools: CT SL HZ AO. Wrote the paper: SKJL CT LG MN AO.