About the Authors

Damien Farrell

Contributed equally to this work with: Damien Farrell, Ronan G. Shaughnessy

Affiliation UCD School of Veterinary Medicine, University College Dublin, Dublin, Ireland

Ronan G. Shaughnessy

Contributed equally to this work with: Damien Farrell, Ronan G. Shaughnessy

Affiliation UCD School of Veterinary Medicine, University College Dublin, Dublin, Ireland

Louise Britton

Affiliation UCD School of Veterinary Medicine, University College Dublin, Dublin, Ireland

David E. MacHugh

Affiliations UCD School of Agriculture and Food Science, University College Dublin, Dublin, Ireland, UCD Conway Institute, University College Dublin, Dublin, Ireland

Bryan Markey

Affiliation UCD School of Veterinary Medicine, University College Dublin, Dublin, Ireland

Stephen V. Gordon

stephen.gordon@ucd.ie

Affiliations UCD School of Veterinary Medicine, University College Dublin, Dublin, Ireland, UCD School of Medicine, University College Dublin, Dublin, Ireland, UCD School of Biomolecular and Biomedical Science, University College Dublin, Dublin, Ireland, UCD Conway Institute, University College Dublin, Dublin, Ireland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RS LB DEM BM SVG. Performed the experiments: RS LB BM. Analyzed the data: DF RS. Wrote the paper: DF RS SVG.