About the Authors

Ranjeet Kumar

Current address: Laboratory of Biochemistry and Molecular Biology, Tomsk State University, Tomsk, Russian Federation

Affiliation School of Science and Technology, The Life Science Center-Biology, Örebro University, Örebro, Sweden

Ajay Pradhan

Affiliation School of Science and Technology, The Life Science Center-Biology, Örebro University, Örebro, Sweden

Faisal Ahmad Khan

Affiliation School of Science and Technology, The Life Science Center-Biology, Örebro University, Örebro, Sweden

Pia Lindström

Affiliation Boliden Mineral AB, Boliden, Sweden

Daniel Ragnvaldsson

Affiliation Envix Nord AB, Umeå, Sweden

Per Ivarsson

Affiliation ALS Scandinavia AB, Täby, Sweden

Per-Erik Olsson

Affiliation School of Science and Technology, The Life Science Center-Biology, Örebro University, Örebro, Sweden

Jana Jass

jana.jass@oru.se

Affiliation School of Science and Technology, The Life Science Center-Biology, Örebro University, Örebro, Sweden

Competing Interests

Boliden Mineral AB, Envix Nord AB and ALS Scandinavia AB provided support in the form of salaries for authors PL, DR and PI, respectively, but did not have any additional role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. The specific roles of these authors are articulated in the ‘author contributions’ section. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RK PL DR PI PEO JJ. Performed the experiments: RK AP FAK. Analyzed the data: RK AP FAK PEO JJ. Contributed reagents/materials/analysis tools: PL DR PI PEO JJ. Wrote the paper: RK AP FAK PL DR PI PEO JJ.