About the Authors

Clara Escudero Duch

‡ These authors are joint first authors on this work.

Affiliation Department of Microbiology III, Faculty of Biological Sciences, Universidad Complutense de Madrid, Madrid, Spain

Richard A. J. Williams

‡ These authors are joint first authors on this work.

Affiliations Department of Zoology and Physical Anthropology, Faculty of Biological Sciences, Universidad Complutense de Madrid, Madrid, Spain, Natural Sciences, Saint Louis University, Madrid, Spain, Department of Ecology and Evolutionary Biology & Natural History Museum, University of Kansas, Lawrence, Kansas 66045, United States of America

Robert M. Timm

Affiliation Department of Ecology and Evolutionary Biology & Natural History Museum, University of Kansas, Lawrence, Kansas 66045, United States of America

Javier Perez-Tris

Affiliation Department of Zoology and Physical Anthropology, Faculty of Biological Sciences, Universidad Complutense de Madrid, Madrid, Spain

Laura Benitez

lbenitez@bio.ucm.es

Affiliation Department of Microbiology III, Faculty of Biological Sciences, Universidad Complutense de Madrid, Madrid, Spain

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RAJW RMT LB. Performed the experiments: CE LB. Analyzed the data: RAJW JPT LB. Contributed reagents/materials/analysis tools: JPT LB. Wrote the paper: CE RAJW RMT JPT LB. Collected samples: RAJW RMT.