About the Authors

Adam G. Hart

Contributed equally to this work with: Adam G. Hart, Richard N. Rolfe, Anne E. Goodenough

ahart@glos.ac.uk

Affiliation School of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Richard N. Rolfe

Contributed equally to this work with: Adam G. Hart, Richard N. Rolfe, Anne E. Goodenough

Affiliation School of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Shantelle Dandy

‡ These authors also contributed equally to this work.

Affiliation School of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Hannah Stubbs

‡ These authors also contributed equally to this work.

Affiliation School of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Dougal MacTavish

‡ These authors also contributed equally to this work.

Affiliation Mankwe Wildlife Reserve, Rustenburg, Northwest Province, South Africa

Lynne MacTavish

‡ These authors also contributed equally to this work.

Affiliation Mankwe Wildlife Reserve, Rustenburg, Northwest Province, South Africa

Anne E. Goodenough

Contributed equally to this work with: Adam G. Hart, Richard N. Rolfe, Anne E. Goodenough

Affiliation School of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Competing Interests

Co-authors Dougal MacTavish and Lynne MacTavish are employed by Mankwe Wildlife Reserve. There are no patents, products in development, or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AGH AEG RNR HS. Performed the experiments: AGH AEG RNR SD LM DM. Analyzed the data: AGH AEG RNR SD. Contributed reagents/materials/analysis tools: HS. Wrote the paper: AGH AEG RNR SD HS LM DM.