About the Authors

Tara Grinnage-Pulley

Current address: Department of Epidemiology, College of Public Health, University of Iowa, Iowa City, Iowa, United States of America

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Qijing Zhang

zhang123@iastate.edu

Affiliation Department of Veterinary Microbiology and Preventive Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Competing Interests

Qijing Zhang is a PLOS ONE Editorial Board member, and Tara Grinnage-Pulley received commercial funding from Fort Dodge Animal Health and Pfizer Animal Health. However, these do not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TGP QZ. Performed the experiments: TGP. Analyzed the data: TGP QZ. Contributed reagents/materials/analysis tools: QZ. Wrote the paper: TGP QZ.