About the Authors

Nicholas A. Fisichelli

nicholas_fisichelli@nps.gov

Affiliation Natural Resource Science and Stewardship, US National Park Service, Fort Collins, Colorado, United States of America

Gregor W. Schuurman

Affiliation Natural Resource Science and Stewardship, US National Park Service, Fort Collins, Colorado, United States of America

William B. Monahan

Affiliation Natural Resource Science and Stewardship, US National Park Service, Fort Collins, Colorado, United States of America

Pamela S. Ziesler

Affiliation Natural Resource Science and Stewardship, US National Park Service, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NAF GWS WBM. Performed the experiments: NAF GWS WBM. Analyzed the data: NAF GWS WMB PSZ. Wrote the paper: NAF GWS WMB PSZ.