About the Authors

Lu Hou

Affiliations Key Laboratory of Agricultural Integrated Pest Management, Institute of Plant Protection, Qinghai Academy of Agriculture and Forestry Sciences, Xining, Qinghai, China, Department of Plant Pathology, Washington State University, Pullman, Washington, United States of America, State Key Laboratory of Crop Stress Biology in Arid Areas and College of Plant Protection, Northwest A&F University, Yangling, Shaanxi, China

Xianming Chen

xianming@wsu.edu

Affiliations Department of Plant Pathology, Washington State University, Pullman, Washington, United States of America, US Department of Agriculture, Agricultural Research Service, Wheat Genetics, Quality, Physiology and Disease Research Unit, Pullman, Washington, United States of America

Meinan Wang

Affiliation Department of Plant Pathology, Washington State University, Pullman, Washington, United States of America

Deven R. See

Affiliations Department of Plant Pathology, Washington State University, Pullman, Washington, United States of America, US Department of Agriculture, Agricultural Research Service, Wheat Genetics, Quality, Physiology and Disease Research Unit, Pullman, Washington, United States of America

Shiaoman Chao

Affiliation US Department of Agriculture, Agricultural Research Service, Cereal Crops Research, Fargo, North Dakota, United States of America

Peter Bulli

Affiliation Department of Crop and Soil Sciences, Washington State University, Pullman, Washington, United States of America

Jinxue Jing

Affiliation State Key Laboratory of Crop Stress Biology in Arid Areas and College of Plant Protection, Northwest A&F University, Yangling, Shaanxi, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XMC. Performed the experiments: LH MNW. Analyzed the data: LH XMC MNW PB. Contributed reagents/materials/analysis tools: XMC DRS SMC JXJ. Wrote the paper: LH XMC. Revised the manuscript: MNW DRS PB SMC.