About the Authors

Takashi A. Inoue

Contributed equally to this work with: Takashi A. Inoue, Tetsuo Ito, Hiroshi Hagiya, Tamako Hata, Kiyoshi Asaoka, Fumio Yokohari, Kinuko Niihara

inoueatp@affrc.go.jp

Affiliation Japanese National Institute of Agrobiological Sciences, Ôwashi 1–2, Tsukuba, Ibaraki, 305–8634, Japan

Tetsuo Ito

Contributed equally to this work with: Takashi A. Inoue, Tetsuo Ito, Hiroshi Hagiya, Tamako Hata, Kiyoshi Asaoka, Fumio Yokohari, Kinuko Niihara

Affiliation Kibougaoka 34–19, Asahi, Yokohama, Kanagawa, 241–0624, Japan

Hiroshi Hagiya

Contributed equally to this work with: Takashi A. Inoue, Tetsuo Ito, Hiroshi Hagiya, Tamako Hata, Kiyoshi Asaoka, Fumio Yokohari, Kinuko Niihara

Affiliation Department of Natural Science, Faculty of Knowledge Engineering, Tokyo City University, Tamatudzumi, Setagaya, Tokyo, 158–8557, Japan

Tamako Hata

Contributed equally to this work with: Takashi A. Inoue, Tetsuo Ito, Hiroshi Hagiya, Tamako Hata, Kiyoshi Asaoka, Fumio Yokohari, Kinuko Niihara

Affiliation Japanese National Institute of Agrobiological Sciences, Ôwashi 1–2, Tsukuba, Ibaraki, 305–8634, Japan

Kiyoshi Asaoka

Contributed equally to this work with: Takashi A. Inoue, Tetsuo Ito, Hiroshi Hagiya, Tamako Hata, Kiyoshi Asaoka, Fumio Yokohari, Kinuko Niihara

Affiliation Japanese National Institute of Agrobiological Sciences, Ôwashi 1–2, Tsukuba, Ibaraki, 305–8634, Japan

Fumio Yokohari

Contributed equally to this work with: Takashi A. Inoue, Tetsuo Ito, Hiroshi Hagiya, Tamako Hata, Kiyoshi Asaoka, Fumio Yokohari, Kinuko Niihara

Affiliation Division of Biology, Department of Earth System Science, Faculty of Science, Fukuoka University, Shichikuma 8-19-1, Jounan, Fukuoka, Fukuoka, 814–0180, Japan

Kinuko Niihara

Contributed equally to this work with: Takashi A. Inoue, Tetsuo Ito, Hiroshi Hagiya, Tamako Hata, Kiyoshi Asaoka, Fumio Yokohari, Kinuko Niihara

Affiliation Department of Natural Science, Faculty of Knowledge Engineering, Tokyo City University, Tamatudzumi, Setagaya, Tokyo, 158–8557, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TAI TI HH KN. Performed the experiments: TAI TI HH KN. Analyzed the data: TAI TI HH KN. Contributed reagents/materials/analysis tools: TAI TI HH KN. Wrote the paper: TAI TH KA FY KN.