About the Authors

Yuji Suzuki

Affiliation Center for Integrated Human Brain Science, Brain Research Institute, University of Niigata, Niigata, Japan

Yukihiro Nakamura

Affiliation Center for Integrated Human Brain Science, Brain Research Institute, University of Niigata, Niigata, Japan

Kenichi Yamada

Affiliation Center for Integrated Human Brain Science, Brain Research Institute, University of Niigata, Niigata, Japan

Hironaka Igarashi

Affiliation Center for Integrated Human Brain Science, Brain Research Institute, University of Niigata, Niigata, Japan

Kensaku Kasuga

Affiliation Department of Molecular Genetics, Brain Research Institute, University of Niigata, Niigata, Japan

Yuichi Yokoyama

Affiliation Department of Psychiatry, Faculty of Medicine, University of Niigata, Niigata, Japan

Takeshi Ikeuchi

Affiliation Department of Molecular Genetics, Brain Research Institute, University of Niigata, Niigata, Japan

Masatoyo Nishizawa

Affiliation Department of Neurology, Brain Research Institute, University of Niigata, Niigata, Japan

Ingrid L. Kwee

Affiliation Department of Neurology, University of California Davis, Davis, California, United States of America

Tsutomu Nakada

tnakada@bri.niigata-u.ac.jp

Affiliations Center for Integrated Human Brain Science, Brain Research Institute, University of Niigata, Niigata, Japan, Department of Neurology, University of California Davis, Davis, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TN. Performed the experiments: YS YN KY KK YY TI. Analyzed the data: YS HI MN ILK TN. Wrote the paper: YS ILK TN.