About the Authors

Yu Sun

Affiliation Centre for Gene Regulation and Expression, College of Life Sciences, University of Dundee, Dundee, United Kingdom

Jacob George

j.george@dundee.ac.uk (JG); s.rocha@dundee.ac.uk (SR)

Affiliation Division of Medical Sciences, Ninewells Hospital and Medical School, Dundee, United Kingdom

Sonia Rocha

j.george@dundee.ac.uk (JG); s.rocha@dundee.ac.uk (SR)

Affiliation Centre for Gene Regulation and Expression, College of Life Sciences, University of Dundee, Dundee, United Kingdom

Competing Interests

The authors confirm that the co-author Sonia Rocha is a PLOS One Editorial board member and that this does not alter the authors' adherence to PLOS One editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: JG SR. Performed the experiments: YS. Analyzed the data: YS SR. Wrote the paper: YS JG SR.