About the Authors

Robert S. Alonso

Affiliations Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America, Western Ecological Research Center, Biological Resources Discipline, United States Geological Survey, Thousand Oaks, California, United States of America

Brett T. McClintock

Affiliation National Marine Mammal Laboratory, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, Seattle, Washington, United States of America

Lisa M. Lyren

Affiliation Western Ecological Research Center, Biological Resources Discipline, United States Geological Survey, Carlsbad, California, United States of America

Erin E. Boydston

Affiliation Western Ecological Research Center, Biological Resources Discipline, United States Geological Survey, Thousand Oaks, California, United States of America

Kevin R. Crooks

kevin.crooks@colostate.edu

Affiliation Department of Fish, Wildlife, and Conservation Biology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

A commercial source, The Irvine Company, provided funding for this research. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RSA BTM LML EEB KRC. Performed the experiments: RSA BTM LML EEB KRC. Analyzed the data: RSA BTM. Contributed reagents/materials/analysis tools: RSA BTM LML EEB KRC. Wrote the paper: RSA BTM LML EEB KRC.