About the Authors

Alexander C. Lees

alexanderlees@btopenworld.com

Affiliation Museu Paraense Emílio Goeldi, Belém, PA, Brazil

Nárgila G. Moura

Contributed equally to this work with: Nárgila G. Moura, Arlete Silva de Almeida

Affiliation Museu Paraense Emílio Goeldi, Belém, PA, Brazil

Arlete Silva de Almeida

Contributed equally to this work with: Nárgila G. Moura, Arlete Silva de Almeida

Affiliations Museu Paraense Emílio Goeldi, Belém, PA, Brazil, Programa de Pós-Graduação em Ciências Ambientais, Universidade Federal do Pará/ Museu Paraense Emílio Goeldi, Belém—Pará, Brazil

Ima C. G. Vieira

Affiliations Museu Paraense Emílio Goeldi, Belém, PA, Brazil, Programa de Pós-Graduação em Ciências Ambientais, Universidade Federal do Pará/ Museu Paraense Emílio Goeldi, Belém—Pará, Brazil

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ACL ICGV. Performed the experiments: ACL NGM ASdA. Analyzed the data: ACL NGM ASdA. Contributed reagents/materials/analysis tools: ASdA. Wrote the paper: ACL ICGV.