About the Authors

Zhongqiang Zhou

Affiliation State Key Laboratory of Ophthalmology and Division of Preventive Ophthalmology, Zhongshan Ophthalmic Center, Sun Yat-sen University, Guangzhou, China

Ian G. Morgan

Affiliations State Key Laboratory of Ophthalmology and Division of Preventive Ophthalmology, Zhongshan Ophthalmic Center, Sun Yat-sen University, Guangzhou, China, ARC Centre of Excellence in Vision Science, Research School of Biology, Australian National University, Canberra, Australia

Qianyun Chen

Affiliation State Key Laboratory of Ophthalmology and Division of Preventive Ophthalmology, Zhongshan Ophthalmic Center, Sun Yat-sen University, Guangzhou, China

Ling Jin

Affiliation State Key Laboratory of Ophthalmology and Division of Preventive Ophthalmology, Zhongshan Ophthalmic Center, Sun Yat-sen University, Guangzhou, China

Mingguang He

Affiliation State Key Laboratory of Ophthalmology and Division of Preventive Ophthalmology, Zhongshan Ophthalmic Center, Sun Yat-sen University, Guangzhou, China

Nathan Congdon

ncongdon1@gmail.com

Affiliations State Key Laboratory of Ophthalmology and Division of Preventive Ophthalmology, Zhongshan Ophthalmic Center, Sun Yat-sen University, Guangzhou, China, ORBIS International, New York, New York, United States of America

Competing Interests

Co-author Mingguang is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: ZZ IGM MH NC. Performed the experiments: ZZ QC MH. Analyzed the data: IGM LJ NC. Contributed reagents/materials/analysis tools: MH. Wrote the paper: NC ZZ IGM. Reviewed and revised the manuscript: ZZ IGM QC LJ MH NC.