About the Authors

Zirong Gu

Affiliation Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

Fumiyasu Imai

Affiliation Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

In Jung Kim

Affiliation Department of Ophthalmology and Visual Science and Department of Neurobiology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Hiroko Fujita

Affiliation RIKEN Brain Science Institute, Saitama, Japan

Kei ichi Katayama

Affiliation Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

Kensaku Mori

Affiliation Department of Physiology, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Yoshihiro Yoshihara

Affiliation RIKEN Brain Science Institute, Saitama, Japan

Yutaka Yoshida

yutaka.yoshida@cchmc.org

Affiliation Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZG FI Y. Yoshida. Performed the experiments: ZG FI HF KK. Analyzed the data: ZG FI Y. Yoshida. Contributed reagents/materials/analysis tools: IJK HF KM Y. Yoshihara. Wrote the paper: Y. Yoshida.