About the Authors

Yoko Inui

inui@cc.osaka-kyoiku.ac.jp

Affiliation Department of Arts and Sciences, Osaka Kyoiku University, Kashiwara, Osaka, Japan

Usun Shimizu-kaya

Affiliation Graduate School of Human and Environmental Studies, Kyoto University, Sakyo-ku, Kyoto, Japan

Tadahiro Okubo

Current address: Kansai University Dai-ichi Senior High School, Suita, Osaka, Japan

Affiliation Graduate School of Human and Environmental Studies, Kyoto University, Sakyo-ku, Kyoto, Japan

Eri Yamsaki

Current address: Institute of Evolutionary Biology and Environmental Studies, University of Zurich, Zurich, Switzerland

Affiliation Center for Ecological Research, Kyoto University, Otsu, Shiga, Japan

Takao Itioka

Affiliation Graduate School of Human and Environmental Studies, Kyoto University, Sakyo-ku, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YI TI. Performed the experiments: YI EY US TO. Analyzed the data: YI. Contributed reagents/materials/analysis tools: YI EY. Wrote the paper: YI TI.