About the Authors

Saioa Fernández-Beaskoetxea

Affiliation Museo Nacional de Ciencias Naturales, CSIC, Madrid, Spain

Luis M. Carrascal

Affiliation Museo Nacional de Ciencias Naturales, CSIC, Madrid, Spain

Andrés Fernández-Loras

Affiliation Museo Nacional de Ciencias Naturales, CSIC, Madrid, Spain

Matthew C. Fisher

matthew.fisher@imperial.ac.uk (MCF); bosch@mncn.cisc.es (JB)

Affiliation Department of Infectious Disease Epidemiology, Imperial College London, St. Mary’s Hospital, London, United Kingdom

Jaime Bosch

matthew.fisher@imperial.ac.uk (MCF); bosch@mncn.cisc.es (JB)

Affiliations Museo Nacional de Ciencias Naturales, CSIC, Madrid, Spain, Centro de Investigación, Seguimiento y Evaluación, Parque Nacional de la Sierra de Guadarrama, Rascafría, Spain

Competing Interests

Funding for this study was provided by Fundación General CSIC and Banco Santander. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JB SFB. Performed the experiments: SFB AFL JB. Analyzed the data: LMC SFB JB. Contributed reagents/materials/analysis tools: SFB JB LMC MCF AFL. Wrote the paper: SFB JB LMC MCF AFL.