About the Authors

Jennifer L. Wilkening

Jennifer.Wilkening@colorado.edu

Affiliation Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, United States of America

Chris Ray

Affiliation Department of Ecology and Evolutionary Biology and Institute for Arctic and Alpine Research, University of Colorado, Boulder, Colorado, United States of America

Johanna Varner

Affiliation Department of Biology, University of Utah, Salt Lake City, Utah, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JLW CR JV. Performed the experiments: JLW JV. Analyzed the data: JLW CR. Contributed reagents/materials/analysis tools: JLW CR. Wrote the paper: JLW CR JV.