About the Authors

Simon P. Rout

Affiliation Department of Biological Sciences, School of Applied Sciences, University of Huddersfield, Queensgate, Huddersfield, HD1 3DH, United Kingdom

Christopher J. Charles

Affiliation Department of Biological Sciences, School of Applied Sciences, University of Huddersfield, Queensgate, Huddersfield, HD1 3DH, United Kingdom

Eva J. Garratt

Affiliation Department of Biological Sciences, School of Applied Sciences, University of Huddersfield, Queensgate, Huddersfield, HD1 3DH, United Kingdom

Andrew P. Laws

Affiliation Department of Biological Sciences, School of Applied Sciences, University of Huddersfield, Queensgate, Huddersfield, HD1 3DH, United Kingdom

John Gunn

Affiliation School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, B15 2TT, United Kingdom

Paul N. Humphreys

p.n.humphreys@hud.ac.uk

Affiliation Department of Biological Sciences, School of Applied Sciences, University of Huddersfield, Queensgate, Huddersfield, HD1 3DH, United Kingdom

Competing Interests

Radioactive Waste Management Limited is a wholly owned subsidiary of the Nuclear Decommissioning Authority, which is an Executive Non-Departmental Public Body of the Department of the Energy and Climate Change of the UK government. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials

Author Contributions

Conceived and designed the experiments: PNH SR CC JG. Performed the experiments: SR CC EG. Analyzed the data: PNH SR CC AL. Wrote the paper: SR PNH AL.