About the Authors

Jacek K. Pijanka

Affiliation Structural Biophysics Group, School of Optometry and Vision Sciences, Cardiff University, United Kingdom

Martin T. Spang

Affiliation Structural Biophysics Group, School of Optometry and Vision Sciences, Cardiff University, United Kingdom

Thomas Sorensen

Affiliation Diamond Light Source, Didcot, United Kingdom

Jun Liu

Affiliation Department of Biomedical Engineering, The Ohio State University, Columbus, Ohio, United States of America

Thao D. Nguyen

Affiliation Department of Mechanical Engineering, Johns Hopkins University, Baltimore, Maryland, United States of America

Harry A. Quigley

Affiliation Glaucoma Center of Excellence, Wilmer Ophthalmological Institute, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Craig Boote

bootec@cardiff.ac.uk

Affiliation Structural Biophysics Group, School of Optometry and Vision Sciences, Cardiff University, United Kingdom

Competing Interests

Craig Boote is an Academic Editor for PLOS ONE. One of the authors (Dr. T. Sorensen) is employed by Diamond Light Source Ltd, where much of the data for the study was collected. There are no patents, products in development or marketed products to declare. The authors can confirm that this does not alter their adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JKP TS CB. Performed the experiments: JKP TS CB. Analyzed the data: JKP MTS JL CB. Contributed reagents/materials/analysis tools: TS TDN JL HAQ. Wrote the paper: JKP JL TDN HAQ CB.