About the Authors

Terry A. Klein

Affiliation Force Health Protection and Preventive Medicine, 65th Medical Brigade/US Army MEDDAC-Korea, Unit 15281, APO AP 96205–528, United States of America

Heung-Chul Kim

Affiliation 5th Medical Detachment, 168th Multifunctional Medical Battalion, 65th Medical Brigade, Unit 15247, APO AP 96205–5247, United States of America

Sung-Tae Chong

Affiliation 5th Medical Detachment, 168th Multifunctional Medical Battalion, 65th Medical Brigade, Unit 15247, APO AP 96205–5247, United States of America

Jeong-Ah Kim

Affiliation Department of Microbiology, College of Medicine, Institute of Biomedical Science & Food Safety, Korea University, 126–1, 5-ga, Anam-dong, Seongbuk-gu, Seoul, 136–705, Republic of Korea

Sook-Young Lee

Affiliation Department of Microbiology, College of Medicine, Institute of Biomedical Science & Food Safety, Korea University, 126–1, 5-ga, Anam-dong, Seongbuk-gu, Seoul, 136–705, Republic of Korea

Won-Keun Kim

Affiliation Department of Microbiology, College of Medicine, Institute of Biomedical Science & Food Safety, Korea University, 126–1, 5-ga, Anam-dong, Seongbuk-gu, Seoul, 136–705, Republic of Korea

Peter V. Nunn

Current address: Navy Entomology Center of Excellence, P.O. Box 43, Bldg 937, Naval Air Station, Jacksonville, FL, 32212–0043, United States of America

Affiliation 5th Medical Detachment, 168th Multifunctional Medical Battalion, 65th Medical Brigade, Unit 15247, APO AP 96205–5247, United States of America

Jin-Won Song

jwsong@korea.ac.kr

Affiliation Department of Microbiology, College of Medicine, Institute of Biomedical Science & Food Safety, Korea University, 126–1, 5-ga, Anam-dong, Seongbuk-gu, Seoul, 136–705, Republic of Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TAK HCK JWS. Performed the experiments: STC JAK SYL WKK. Analyzed the data: TAK HCK JWS. Contributed reagents/materials/analysis tools: TAK JWS. Wrote the paper: TAK HCK JWS. Conducted field surveillance: TAK PVN HCK.