About the Authors

Ryan Mathur

Affiliation Sheffield Institute for Translational Neuroscience, University of Sheffield, Sheffield, England, United Kingdom

Paul G. Ince

Affiliation Sheffield Institute for Translational Neuroscience, University of Sheffield, Sheffield, England, United Kingdom

Thais Minett

Affiliation Institute of Public Health, University of Cambridge, Cambridge, England, United Kingdom

Claire J. Garwood

Affiliation Sheffield Institute for Translational Neuroscience, University of Sheffield, Sheffield, England, United Kingdom

Pamela J. Shaw

Affiliation Sheffield Institute for Translational Neuroscience, University of Sheffield, Sheffield, England, United Kingdom

Fiona E. Matthews

Affiliation MRC Biostatistics Unit, Cambridge, England, United Kingdom

Carol Brayne

Affiliation Institute of Public Health, University of Cambridge, Cambridge, England, United Kingdom

Julie E. Simpson

Contributed equally to this work with: Julie E. Simpson, Stephen B. Wharton

julie.simpson@sheffield.ac.uk

Affiliation Sheffield Institute for Translational Neuroscience, University of Sheffield, Sheffield, England, United Kingdom

Stephen B. Wharton

Contributed equally to this work with: Julie E. Simpson, Stephen B. Wharton

Affiliation Sheffield Institute for Translational Neuroscience, University of Sheffield, Sheffield, England, United Kingdom

on behalf of the MRC Cognitive Function and Ageing Neuropathology Study Group

Membership of the MRC Cognitive Function and Ageing Neuropathology Study Group are listed on the CFAS website (www.cfas.ac.uk/pages/contacts/index.html).

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SBW PGI. Performed the experiments: RM JES. Analyzed the data: TM FM SBW. Wrote the paper: RM JES SBW PGI TM CJG PJS FEM CB.