About the Authors

Daniel L. Morgan

morgan3@niehs.nih.gov

‡ These authors contributed equally to this work.

Affiliation Division of the National Toxicology Program, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

B. Alex Merrick

‡ These authors contributed equally to this work.

Affiliation Division of the National Toxicology Program, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

Kevin E. Gerrish

‡ These authors contributed equally to this work.

Affiliation Division of Intramural Research, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

Patricia S. Stockton

Affiliation Division of the National Toxicology Program, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

Yu Wang

Affiliation Division of the National Toxicology Program, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

Julie F. Foley

Affiliation Division of the National Toxicology Program, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

William M. Gwinn

Affiliation Division of the National Toxicology Program, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

Francine L. Kelly

Affiliation Duke University Medical Center, Durham, North Carolina, United States of America

Scott M. Palmer

Affiliation Duke University Medical Center, Durham, North Carolina, United States of America

Thai-Vu T. Ton

Affiliation Division of the National Toxicology Program, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

Gordon P. Flake

‡ These authors contributed equally to this work.

Affiliation Division of the National Toxicology Program, National Institute of Environmental Health Sciences, Research Triangle Park, North Carolina, United States of America

Competing Interests

The authors have the following interests. These studies were conducted at the NIEHS inhalation facility under contract to Alion Science and Technology, Inc. There are no patents, products in development, or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: DLM GPF BAM. Performed the experiments: KEG PSS JFF WMG FLK TT YW. Analyzed the data: DLM GPF BAM KEG SMP. Contributed reagents/materials/analysis tools: DLM GPF BAM KEG PSS JFF WMG FLK SMP TT YW. Wrote the paper: DLM BAM KEG PSS JFF WMG FLK SMP GPF.