About the Authors

Hannah Arem

Hannah.arem@nih.gov

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Kai Yu

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Xiaoqin Xiong

Affiliation Information Management Systems, Inc., Calverton, Maryland, United States of America

Kristin Moy

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Neal D. Freedman

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Susan T. Mayne

Affiliation Yale School of Public Health/Yale Cancer Center, New Haven, Connecticut, United States of America

Demetrius Albanes

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Alan A. Arslan

Affiliation Departments of Population Health, Obstetrics and Gynecology (Obs/Gyn) and Environmental Medicine, New York University, New York, New York, United States of America

Melissa Austin

Affiliation Department of Epidemiology, University of Washington, Seattle, Washington, United States of America

William R. Bamlet

Affiliation Department of Epidemiology, Mayo Clinic, Rochester, Minnesota, United States of America

Laura Beane-Freeman

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Paige Bracci

Affiliation Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America

Federico Canzian

Affiliation Division of Cancer Epidemiology, German Cancer Research Center (DKFZ), Heidelberg, Germany

Michelle Cotterchio

Affiliation Dalla Lana School of Public Health, University of Toronto; Prevention and Cancer Control, Cancer Care Ontario Toronto, Ontario, Canada

Eric J. Duell

Affiliation Catalan Institute of Oncology (ICO-IDIBELL), Barcelona, Spain

Steve Gallinger

Affiliation Samuel Lunenfeld Research Institute, Mount Sinai Hospital, University of Toronto, Toronto, Canada

Graham G. Giles

Affiliation Cancer Epidemiology Centre, Cancer Council Victoria and Centre for MEGA Epidemiology, School of Population Health, the University of Melbourne, Melbourne, Australia

Michael Goggins

Affiliation Departments of Oncology, Pathology and Medicine, The Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Phyllis J. Goodman

Affiliation Cleveland Clinic, Glickman Urological and Kidney Institute, Cleveland, Ohio, United States of America

Patricia Hartge

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Manal Hassan

Affiliation Department of Gastrointestinal Medical Oncology, The University of Texas M. D. Anderson Cancer Center, Houston, Texas, United States of America

Kathy Helzlsouer

Affiliation MD Mercy, Baltimore, Maryland, United States of America

Brian Henderson

Affiliation Department of Preventative Medicine, School of Medicine, University of Southern California, Los Angeles, California, United States of America

Elizabeth A. Holly

Affiliation Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America

Robert Hoover

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Eric J. Jacobs

Affiliation Epidemiology Research Program, American Cancer Society, Atlanta, Georgia, United States of America

Aruna Kamineni

Affiliation GroupHealth Research Institute, Seattle, Washington, United States of America

Alison Klein

Affiliation MD Mercy, Baltimore, Maryland, United States of America

Eric Klein

Affiliation Cleveland Clinic, Glickman Urological and Kidney Institute, Cleveland, Ohio, United States of America

Laurence N. Kolonel

Affiliation University of Hawaii Cancer Center, Manoa, Hawaii, United States of America

Donghui Li

Affiliation Department of Gastrointestinal Medical Oncology, The University of Texas M. D. Anderson Cancer Center, Houston, Texas, United States of America

Núria Malats

Affiliation Molecular Pathology Program, Spanish National Cancer Research Center, Madrid, Spain

Satu Männistö

Affiliation National Institute for Health and Welfare, Department of Chronic Disease Prevention, Helsinki, Finland

Marjorie L. McCullough

Affiliation Epidemiology Research Program, American Cancer Society, Atlanta, Georgia, United States of America

Sara H. Olson

Affiliation Department of Epidemiology and Biostatistics, Memorial Sloan-Kettering Cancer Center, New York, New York, United States of America

Irene Orlow

Affiliation Department of Epidemiology and Biostatistics, Memorial Sloan-Kettering Cancer Center, New York, New York, United States of America

Ulrike Peters

Affiliation Cleveland Clinic, Glickman Urological and Kidney Institute, Cleveland, Ohio, United States of America

Gloria M. Petersen

Affiliation Department of Epidemiology, Mayo Clinic, Rochester, Minnesota, United States of America

Miquel Porta

Affiliation Hospital del Mar Institute of Medical Research (IMIM), and School of Medicine, Barcelona Spain

Gianluca Severi

Affiliation Cancer Epidemiology Centre, Cancer Council Victoria and Centre for MEGA Epidemiology, School of Population Health, the University of Melbourne, Melbourne, Australia

Xiao-Ou Shu

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, and Vanderbilt-Ingram Cancer Center, Vanderbilt University, Nashville, Tennessee, United States of America

Kala Visvanathan

Affiliation MD Mercy, Baltimore, Maryland, United States of America

Emily White

Affiliation Cleveland Clinic, Glickman Urological and Kidney Institute, Cleveland, Ohio, United States of America

Herbert Yu

Affiliation University of Hawaii Cancer Center, Manoa, Hawaii, United States of America

Anne Zeleniuch-Jacquotte

Affiliation Departments of Population Health, Obstetrics and Gynecology (Obs/Gyn) and Environmental Medicine, New York University, New York, New York, United States of America

Wei Zheng

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, and Vanderbilt-Ingram Cancer Center, Vanderbilt University, Nashville, Tennessee, United States of America

Geoffrey S. Tobias

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Dennis Maeder

Affiliation Cancer Genomics Research Laboratory, National Cancer Institute, Division of Cancer Epidemiology and Genetics, Leidos Biomedical Research, Inc., Frederick National Laboratory for Cancer Research, Frederick, Maryland, United States of America

Michelle Brotzman

Affiliation Westat, Rockville, Maryland, United States of America

Harvey Risch

Affiliation Yale School of Public Health/Yale Cancer Center, New Haven, Connecticut, United States of America

Joshua N. Sampson

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Rachael Z. Stolzenberg-Solomon

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, Bethesda, Maryland, United States of America

Competing Interests

Co-authors Xiaoqin Xiong (Information Management Systems, Inc.), Dennis Maeder (Leidos Biomedical Research, Inc.) and Michelle Brotzman (Westat) are employed by commercial enterprises. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HA DM KY RZSS. Performed the experiments: HA XX. Contributed reagents/materials/analysis tools: DDA AAA MA WRB LBF PB FC MC EJD SG GGG MG PJG PH MH KH BH EAH RH EJJ A. Klein A. Kamineni EK LNK DL NM SM MLM SHO IO UP GMP MP GS XS KV EW HY AZJ WZ GST MB HR. Wrote the paper: HA STM JNS HR NDF KM RZSS.