About the Authors

Qing Zeng

Affiliation School of Nature Conservation, Beijing Forestry University, Beijing, China

Yamian Zhang

Affiliation School of Nature Conservation, Beijing Forestry University, Beijing, China

Gongqi Sun

Affiliation School of Nature Conservation, Beijing Forestry University, Beijing, China

Hairui Duo

Affiliation School of Nature Conservation, Beijing Forestry University, Beijing, China

Li Wen

Affiliations School of Nature Conservation, Beijing Forestry University, Beijing, China, Office of Environment and Heritage, Sydney, Australia

Guangchun Lei

guangchun8099@gmail.com

Affiliation School of Nature Conservation, Beijing Forestry University, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: QZ GCL. Performed the experiments: QZ YMZ GQS. Analyzed the data: QZ LW. Contributed reagents/materials/analysis tools: QZ YMZ HRD LW. Wrote the paper: QZ LW GUL.