About the Authors

Richard Kelwick

p.freemont@imperial.ac.uk (PF); r.kitney@imperial.ac.uk (R. Kitney); r.kelwick@imperial.ac.uk (R. Kelwick)

‡ These authors are equal first authors on this work.

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Medicine, South Kensington Campus, London, United Kingdom

Margarita Kopniczky

‡ These authors are equal first authors on this work.

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Medicine, South Kensington Campus, London, United Kingdom

Iain Bower

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Life Sciences, South Kensington Campus, London, United Kingdom

Wenqiang Chi

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Bioengineering, Imperial College London, South Kensington Campus, London, United Kingdom

Matthew Ho Wai Chin

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Bioengineering, Imperial College London, South Kensington Campus, London, United Kingdom

Sisi Fan

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Life Sciences, South Kensington Campus, London, United Kingdom

Jemma Pilcher

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Life Sciences, South Kensington Campus, London, United Kingdom

James Strutt

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Life Sciences, South Kensington Campus, London, United Kingdom

Alexander J. Webb

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Medicine, South Kensington Campus, London, United Kingdom

Kirsten Jensen

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Medicine, South Kensington Campus, London, United Kingdom

Guy-Bart Stan

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Bioengineering, Imperial College London, South Kensington Campus, London, United Kingdom

Richard Kitney

p.freemont@imperial.ac.uk (PF); r.kitney@imperial.ac.uk (R. Kitney); r.kelwick@imperial.ac.uk (R. Kelwick)

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Bioengineering, Imperial College London, South Kensington Campus, London, United Kingdom

Paul Freemont

p.freemont@imperial.ac.uk (PF); r.kitney@imperial.ac.uk (R. Kitney); r.kelwick@imperial.ac.uk (R. Kelwick)

Affiliations Centre for Synthetic Biology and Innovation, South Kensington Campus, London, United Kingdom, Department of Medicine, South Kensington Campus, London, United Kingdom

Competing Interests

The authors declare no conflict of interest and no competing financial interests.

Author Contributions

Conceived and designed the experiments: R. Kelwick MK IB WC MHWC SF JP JS AJW KJ GBS R. Kitney PF. Performed the experiments: R Kelwick MK IB WC MHWC SF JP JS AJW KJ GBS. Analyzed the data: R. Kelwick MK IB WC MHWC SF JP JS AJW KJ GBS R. Kitney PF. Contributed reagents/materials/analysis tools: R. Kelwick MK IB WC MHWC SF JP JS AJW KJ GBS R. Kitney PF. Wrote the paper: R. Kelwick MK IB WC MHWC SF JP JS AJW KJ GBS R. Kitney PF. Supervised and co-ordinated this study: AJW KJ GBS R. Kitney PF.