About the Authors

Chikayo Yamane

Contributed equally to this work with: Chikayo Yamane, Tomohiro Yamazaki

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan

Tomohiro Yamazaki

Contributed equally to this work with: Chikayo Yamane, Tomohiro Yamazaki

Affiliations Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan, Research Fellow of Japan Society for the Promotion of Science, Kojimachi Business Center Building, 5-3-1 Kojimachi, Chiyoda-ku, Tokyo, Japan

Shinji Nakamura

Affiliation Division of Biomedical Imaging Research, Juntendo University Graduate School of Medicine, 2-1-1 Hongo, Bunkyo-ku, Tokyo, Japan

Junji Matsuo

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan

Kasumi Ishida

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan

Sumire Yamazaki

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan

Satoshi Oguri

Affiliations Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan, Hokkaido University Hospital, North-14, West-5, Kita-ku, Sapporo, Japan

Natsumi Shouji

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan

Yasuhiro Hayashi

Affiliations Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan, Hokkaido University Hospital, North-14, West-5, Kita-ku, Sapporo, Japan

Mitsutaka Yoshida

Affiliation Division of Ultrastructural Research, Juntendo University Graduate School of Medicine, 2-1-1 Hongo, Bunkyo-ku, Tokyo, Japan

Yimin

Affiliation Department of Advanced Medicine, Graduate School of Medicine, Hokkaido University, Nishi-7, Kita-15, Kita-ku, Sapporo, Japan

Hiroyuki Yamaguchi

hiroyuki@med.hokudai.ac.jp

Affiliation Department of Medical Laboratory Science, Faculty of Health Sciences, Hokkaido University, North-12, West-5, Kita-ku, Sapporo, Japan

Competing Interests

The authors declare that they have no competing interests.

Author Contributions

Conceived and designed the experiments: HY. Performed the experiments: CY TY NS YH KI SO SY JM. Analyzed the data: SN MY JM HY. Wrote the paper: HY. Critical advice for TEM experiment: MY. Critical advice for in vitro experiment: Y.