About the Authors

Adriano R. Lameira

adriano@orangutan.nl

Affiliations Institute for Biodiversity and Ecosystem Dynamics, University of Amsterdam, Amsterdam, the Netherlands, Pongo Foundation, Oudewater, the Netherlands

Madeleine E. Hardus

Affiliations Institute for Biodiversity and Ecosystem Dynamics, University of Amsterdam, Amsterdam, the Netherlands, Pongo Foundation, Oudewater, the Netherlands

Adrian M. Bartlett

Affiliation Neuroscience Institute, Department of Psychology, Princeton University, Princeton, New Jersey, United States of America

Robert W. Shumaker

Affiliations Indianapolis Zoo, Indianapolis, Indiana, United States of America, Anthropology Department, Indiana University, Bloomington, Indiana, United States of America

Serge A. Wich

Affiliations Institute for Biodiversity and Ecosystem Dynamics, University of Amsterdam, Amsterdam, the Netherlands, Research Centre in Evolutionary Anthropology and Palaeoecology, School of Natural Sciences and Psychology, Liverpool John Moores University, Liverpool, United Kingdom

Steph B. J. Menken

Affiliation Institute for Biodiversity and Ecosystem Dynamics, University of Amsterdam, Amsterdam, the Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ARL MEH. Performed the experiments: ARL MEH. Analyzed the data: ARL AMB. Contributed reagents/materials/analysis tools: ARL MEH AMB. Wrote the paper: ARL MEH AMB RWS SAW SBJM.