About the Authors

Shifa Yang

‡ These authors contributed equally to this work.

Affiliation Shandong Provincial Key Laboratory of Animal Biotechnology and Disease Control and Prevention, College of Animal Science and Technology, Shandong Agricultural University, Shandong, Taian, 271018, PR China

Kai Wei

‡ These authors contributed equally to this work.

Affiliation Shandong Provincial Key Laboratory of Animal Biotechnology and Disease Control and Prevention, College of Animal Science and Technology, Shandong Agricultural University, Shandong, Taian, 271018, PR China

Fengjuan Jia

Affiliation Shandong Provincial Key Laboratory of Animal Biotechnology and Disease Control and Prevention, College of Animal Science and Technology, Shandong Agricultural University, Shandong, Taian, 271018, PR China

Xue Zhao

Affiliation Shandong Provincial Key Laboratory of Animal Biotechnology and Disease Control and Prevention, College of Animal Science and Technology, Shandong Agricultural University, Shandong, Taian, 271018, PR China

Guolin Cui

Affiliation Shandong Provincial Key Laboratory of Animal Biotechnology and Disease Control and Prevention, College of Animal Science and Technology, Shandong Agricultural University, Shandong, Taian, 271018, PR China

Fanxia Guo

Affiliation Shandong Provincial Key Laboratory of Animal Biotechnology and Disease Control and Prevention, College of Animal Science and Technology, Shandong Agricultural University, Shandong, Taian, 271018, PR China

Ruiliang Zhu

zhurl1963@163.com

Affiliation Shandong Provincial Key Laboratory of Animal Biotechnology and Disease Control and Prevention, College of Animal Science and Technology, Shandong Agricultural University, Shandong, Taian, 271018, PR China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RZ SY KW. Performed the experiments: SY KW FJ XZ GC FG. Analyzed the data: RZ SY KW. Contributed reagents/materials/analysis tools: RZ SY FJ KW. Wrote the paper: SY KW FJ RZ.