About the Authors

Aftab Ahmad

Current address: Department of Biochemistry, Center of Advanced Studies (CAS), University of Agriculture, Faisalabad, 38040, Pakistan

Affiliation Laboratory of Plant Molecular Improvement, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Yasuo Niwa

Affiliation Laboratory of Plant Molecular Improvement, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Shingo Goto

Current address: Okitsu Citrus Research Station, NARO Institute of Fruit Tree Science, National Agriculture and Food Research Organization, Shimizu, Shizuoka 424–0292, Japan

Affiliation Laboratory of Plant Molecular Improvement, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Kyoko Kobayashi

Affiliation Laboratory of Plant Molecular Improvement, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Masanori Shimizu

Current address: School of Health Promotional Science, Tokoha University, 1230 Miyakoda, Hamamatsu 431–2102, Japan

Affiliation Laboratory of Plant Molecular Improvement, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Sohei Ito

Affiliation Laboratory of Protein Engineering, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Yumiko Usui

Affiliation Laboratory of Molecular Fooineering, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Tsutomu Nakayama

Current Address: Department of Food Science and Technology, Nippon Veterinary and Life Science University, Tokyo 180–8602, Japan

Affiliation Laboratory of Molecular Fooineering, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Hirokazu Kobayashi

hirokoba@u-shizuoka-ken.ac.jp

Affiliation Laboratory of Plant Molecular Improvement, Graduate School of Nutritional and Environmental Sciences, University of Shizuoka, 52–1 Yada, Suruga, Shizuoka 422–8526, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AA YN SG HK. Performed the experiments: AA. Contributed reagents/materials/analysis tools: KK MS SI YU TN. Wrote the paper: AA HK.