About the Authors

John W. Steinke

js3ch@virginia.edu

Affiliations Department of Medicine, University of Virginia Health System, Charlottesville, Virginia, United States of America, Carter Immunology Center, University of Virginia, Charlottesville, Virginia, United States of America

Lixia Liu

Affiliation Department of Medicine, University of Virginia Health System, Charlottesville, Virginia, United States of America

Ronald B. Turner

Affiliation Department of Pediatrics, University of Virginia Health System, Charlottesville, Virginia, United States of America

Thomas J. Braciale

Affiliations Carter Immunology Center, University of Virginia, Charlottesville, Virginia, United States of America, Department of Pathology, University of Virginia Health System, Charlottesville, Virginia, United States of America

Larry Borish

Affiliations Department of Medicine, University of Virginia Health System, Charlottesville, Virginia, United States of America, Carter Immunology Center, University of Virginia, Charlottesville, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JWS RT TJB LB. Performed the experiments: RT LL JWS. Analyzed the data: JWS TJB LL LB. Contributed reagents/materials/analysis tools: JWS TJB LL LB RT. Wrote the paper: JWS TJB RT LB.