About the Authors

Changjiang Yu

Contributed equally to this work with: Changjiang Yu, Changjiang Sun

Affiliation Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, 266101, PR China

Changjiang Sun

Contributed equally to this work with: Changjiang Yu, Changjiang Sun

Affiliation Beijing Risun Chemical Industry Technology Research institute Co. Ltd, Beijing, 100070, China

Li Yu

Affiliation Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, 266101, PR China

Ming Zhu

Affiliation Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, 266101, PR China

Hua Xu

Affiliation Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, 266101, PR China

Jinshan Zhao

Affiliation Qingdao Institute of Animal Sciences, Qingdao, 266100, China

Yubin Ma

mayb@qibebt.ac.cn (YM); zhougk@qibebt.ac.cn (GZ)

Affiliation Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, 266101, PR China

Gongke Zhou

mayb@qibebt.ac.cn (YM); zhougk@qibebt.ac.cn (GZ)

Affiliation Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao, 266101, PR China

Competing Interests

CS was employed by China Risun Coal Chemicals Group Limited during the course of the study. This study was funded in part by China Risun Coal Chemicals Group Limited. CS is currently employed by Beijing Risun Chemical Industry Technology Research institute Co. Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YC ZG. Performed the experiments: YC ZM SC. Analyzed the data: YC MY XH. Contributed reagents/materials/analysis tools: YC ZM ZJ. Wrote the paper: YC MY ZG.